- BSA, MSc, University of Manitoba
- PhD, University of Oklahoma
- Post-doctoral training: North Carolina State University and University of Minnesota
Loewen ME, Bekar LK, Walz W, Forsyth GW, Gabriel SE. 2004. "pCLCA1 lacks inherent chloride channel activity in an epithelial colon carcinoma cell line." Am J Physiol Gastrointest Liver Physiol, 287: G33-G41.
Loewen ME, Smith NS, Hamilton DL, Grahn BH, Forsyth GW. 2003. "CLCA protein and canine retinal pigment epithelial chloride transport." Am J Physiol Cell Physiol, 285: C1314-C1321.